The Panda’s Thumb Read Free

constrained by history, follows perforce. No one understood this better than Darwin. Ernst Mayr has shown how Darwin, in defending evolution, consistently turned to organic parts and geographic distributions that make the least sense. Which brings me to the giant panda and its “thumb.”
    Giant pandas are peculiar bears, members of the order Carnivora. Conventional bears are the most omnivorous representatives of their order, but pandas have restricted this catholicity of taste in the other direction—they belie the name of their order by subsisting almost entirely on bamboo. They live in dense forests of bamboo at high elevations in the mountains of western China. There they sit, largely unthreatened by predators, munching bamboo ten to twelve hours each day.
    As a childhood fan of Andy Panda, and former owner of a stuffed toy won by some fluke when all the milk bottles actually tumbled at the county fair, I was delighted when the first fruits of our thaw with China went beyond ping pong to the shipment of two pandas to the Washington zoo. I went and watched in appropriate awe. They yawned, stretched, and ambled a bit, but they spent nearly all their time feeding on their beloved bamboo. They sat upright and manipulated the stalks with their forepaws, shedding the leaves and consuming only the shoots.
    I was amazed by their dexterity and wondered how the scion of a stock adapted for running could use its hands so adroitly. They held the stalks of bamboo in their paws and stripped off the leaves by passing the stalks between an apparently flexible thumb and the remaining fingers. This puzzled me. I had learned that a dexterous, opposable thumb stood among the hallmarks of human success. We had maintained, even exaggerated, this important flexibility of our primate forebears, while most mammals had sacrificed it in specializing their digits. Carnivores run, stab, and scratch. My cat may manipulate me psychologically, but he’ll never type or play the piano.
    So I counted the panda’s other digits and received an even greater surprise: there were five, not four. Was the “thumb” a separately evolved sixth finger? Fortunately, the giant panda has its bible, a monograph by D. Dwight Davis, late curator of vertebrate anatomy at Chicago’s Field Museum of Natural History. It is probably the greatest work of modern evolutionary comparative anatomy, and it contains more than anyone would ever want to know about pandas. Davis had the answer, of course.
    The panda’s “thumb” is not, anatomically, a finger at all. It is constructed from a bone called the radial sesamoid, normally a small component of the wrist. In pandas, the radial sesamoid is greatly enlarged and elongated until it almost equals the metapodial bones of the true digits in length. The radial sesamoid underlies a pad on the panda’s forepaw; the five digits form the framework of another pad, the palmar. A shallow furrow separates the two pads and serves as a channelway for bamboo stalks.

    D. L. CRAMER
    The panda’s thumb comes equipped not only with a bone to give it strength but also with muscles to sustain its agility. These muscles, like the radial sesamoid bone itself, did not arise de novo . Like the parts of Darwin’s orchids, they are familiar bits of anatomy remodeled for a new function. The abductor of the radial sesamoid (the muscle that pulls it away from the true digits) bears the formidable name abductor pollicis longus (“the long abductor of the thumb” —pollicis is the genitive of pollex , Latin for “thumb”). Its name is a giveaway. In other carnivores, this muscle attaches to the first digit, or true thumb. Two shorter muscles run between the radial sesamoid and the pollex. They pull the sesamoid “thumb” towards the true digits.
    Does the anatomy of other carnivores give us any clue to the origin of this odd arrangement in pandas? Davis points out that ordinary